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The PDE11A gene occupies >300 kb of genomic DNA and contains 23 exons of generally small size. The four PDE11A variants have different amino termini due to separate transcriptional start sites (Hetman et al., 2000a; Yuasa et al., 2000). Only PDE11A4 contains two complete GAF domains. Oddly enough, the separate transcriptional start sites for PDE11A1-3 result in production of proteins with incomplete GAF domains (Hetman et al., 2000). PDE11A1 completely lacks the N-terminal-most GAF domain, but contains the complete second GAF domain. PDE11A3 and PDE11A2 have one complete GAF domain and one partial GAF domain. The functional consequence(s) of these truncations is unknown, but the truncations do seem to affect catalytic activity.
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